Hi @Tim Graber
A few things to check:
Is the DNS Service on the DCs started
If you use nslookup on one of PC pointing at each DC can you resolve local domain names and external names
If you use nslookup on the DCs pointing at the 8.8.8.8 are you able to resolve external names
If you check in the DNS console, on the server properties are the root hints populated and if you have a Forwarder configured.
Using nslookup on the DC can you resolve external names pointing at one of the root hints and one of the forwards
Gary.
nslookup
server <ip address of DC>
<dc_name>
server 8.8.8.8
google.com
server <ip address of root hint server>
microsoft.com
server <ip address of forwarder>
amazon.com